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They will fire as part of the lifecycle tracking. For this to work you will need to have lifecycle enabled:

Android : Core Implementation and Lifecycle

IOS: Core Implementation and Lifecycle

This is how lifecycle metrics are triggered and why:

IOS: Lifecycle Metrics https://marketing.adobe.com/resources/help/en_US/mobile/ios/metrics.html

Android: Lifecycle Metrics

This is how crashes are handled in Android: Track App Crashes

This is how crashes are handle in IOS : Track App Crashes

Lifecycle tracking implementation is required by a lot of mobile features like link acquisition as well.